Build first, raise later, keep control always
The algorithm walks the tree recursively. At each node, it checks: does this node's bounding box overlap with the query rectangle? If not, the entire subtree gets pruned (skipped). If it does overlap, it tests the node's points against the query and recurses into the children.
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One thing that Nava has got me thinking about is the deep history of gesture. Gesture exists on the borderland between nature and nurture. Certain physical actions are so universal that they communicate effectively not just across human societies but across species: bared teeth, for instance. Other gestures encode more specific cultural patterns, but do so in a way that is more expansive than language.
On this macOS system, that produces 230 fonts across five categories:
The rest of my performance wins were more typical - small reductions in CPU cycles by staring at lots of performance traces.